The posterior part of the base behind the foramen magnum is formed by the supra-occipital part of the occipital bone, so that all the four parts of the bone, which are separate up to the third year, help in the formation of that large opening. Between the foramen magnum and the external occipital protuberance and superior curved line already noticed, the bone attaches the deep muscles of the neck.
and of the sutures between them. In the mid line is a shallow antero-posterior groove for the superior longitudinal blood sinus, and on each side of this irregular depressions are often seen for the Pacchionian bodies (see BRAIN). The base is divided into three fossae, anterior, middle and posterior, each being behind and on a lower level than the one in front of it.
The anterior cranial fossa is formed by the cribriform plate of the ethmoid, near the mid line, freely perforated for the passage of the olfactory nerves. In the mid line, near the front, is a triangular plate rising up which attaches the falx cerebri (see BRAIN) and is called the crista galli. On each side of this is the nasal slit for the nasal branch of the first division of the fifth nerve. On each side of the cribriform plate is the orbital plate of the frontal, while the back part of the fossa has for its floor the body of the sphenoid (pre-sphenoid) near the mid line and the lesser wing (orbito-sphenoid) on each side. Each lesser wing is prolonged back into a tongue-like process, the anterior clinoid process, just internal to which is the optic foramen, and the two foramina are joined by the optic groove for the optic commissure. Behind this groove is a transverse elevation, the olivary eminence, which marks the junction of the pre- and basi-sphenoid parts of the body of the sphenoid bone.
The middle cranial fossa is like an hour-glass placed trans versely, as there is a central constricted, and two lateral ex panded, parts. The central part forms the pituitary fossa for the pituitary body (see BRAIN) and is bounded behind by the wall like dorsum sellae, at the sides of which are the posterior clinoid processes. The olivary eminence, pituitary fossa and dorsum sellae together resemble a Turkish saddle and are often called the sella turcica. The lateral expanded part of the middle cranial fossa is bounded in front by the great wing of the sphenoid (alisphenoid), behind by the front of the petrous part of the tem poral (periotic) and laterally by the squamous part of the tem poral (squamosal). Between the alisphenoid and orbitosphenoid is the sphenoidal fissure already noticed in the orbit, and a little behind this, piercing the alisphenoid, is the posterior opening of the foramen rotundum, through which the second division of the fifth nerve passes into the spheno-maxillary fossa. Further back
the alisphenoid is pierced by the foramen ovale and foramen spinosum, both of which have been already noticed. From the latter a groove for the middle meningeal artery runs forward and outward, and soon divides into anterior and posterior branches, the former of which deepens into a tunnel near the pterion. At the apex of the petrous bone and at the side of the dorsum sellae is the middle lacerated foramen, already noticed, and running inward to this from an aperture in the petrous bone is a groove for the great superficial petrosal nerve which is over laid by the Gasserian ganglion of the fifth nerve.
The posterior cranial fossa is pentagonal in outline, having an anterior border formed by the dorsum sellae, two antero-lateral borders, by the upper borders of the petrous bones, and two postero-lateral curved borders, by the grooves for the lateral sinuses. In the middle of this fossa is the foramen magnum. In front of the foramen magnum the floor of the fossa is formed by the basi-occipital and basi-sphenoid bones, which unite soon after twenty and form a steep slope, downward and backward, known as the clivus. This is slightly grooved from side to side, and lodges the pons and medulla (see BRAIN) and the basilar artery.
On each side of the basi-occipital the posterior surface of the petrous bone bounds the fossa, and lying over the suture between them is the groove for the inferior petrosal venous sinus which leads backward and outward to the jugular foramen already no ticed. About the middle of the posterior surface of the petrous bone is the internal auditory meatus, through which pass the facial and auditory nerves, the pars intermedia (see NERVES : Cranial) and the auditory artery. Close to the antero-lateral part of the foramen magnum is the inner opening of the anterior condylar foramen which is sometimes double for the two bundles of the hypoglossal nerve, and a little in front of and outside this is a heaping up of hone called the tuberculum jugulare, which marks the union of the basi- and ex-occipital bones. The hindmost limit of the posterior fossa in the mid line is marked by an elevation called the internal occipital protuberance, and at this point the grooves for the superior longitudinal, and two lateral sinuses join to form the torcular Herophili (see VEINS). Running from the internal occipital protuberance toward the foramen magnum in the mid line is the internal occipital crest, which attaches the falx cerebelli (see BRAIN) and on each side of this is the cere bellar fossa.