The fundamental difference between softwoods and hardwoods is best understood if we imagine a number of the wider "water fibres" very greatly widened and strung end to end with their terminal walls absorbed, so that long continuous tubes, some times yards in length, are produced. These water-conducting tubes are termed wood-vessels: in cross-section they are often visible to the naked eye as "pores." All commercial hardwoods have wood-vessels, which in the sap-wood contain air with or without liquid sap. In addition to the vessels, the different hard woods have "fibres" of various forms.
Finally medullary rays are present, and in many hardwoods some of them are much thicker than in softwoods and visible to the naked eye in cross-section, producing the so-called "silver grain" on the radial side view of the wood of, for instance, oaks, beech, plane, etc. Their contents in the sapwood are identical with those of softwoods.
branches) runs in a spiral direction as if the trunk had been twisted round its long axis : the grain is then described as spiral or torse, and the timber when cut up is inevitably cross-grained. There are yet other, especially tropical, woods in which the grain more or less swings from a left-handed to a right-handed spiral direction, so that the wood when cut into plain boards shows a double cross-grain or interlocked grain, and when cut along the radii of the trunk ("on the quarter," rift-sawn) shows roe-figure, for instance in mahogany.
In addition to these deviations from the straight grain the structural elements of the wood may pursue a wavy or sinuous course, for instance in so-called rammy-ash. When the waviness is exaggerated the grain is said to be curly.
Knots, being the basal parts of branches that have become embedded in the thickening trunk, are naturally associated with a change of direction of the fibres. The wood of large excrescences on the trunk of certain trees known as burrs exhibits structure similar to that which would be produced by many crowded branches : and a somewhat similar structure is shown by bird's eye maple.
The fibres of most timbers overlap and dovetail with one another at their ends, which are at different levels, and the medul lary rays as seen in circumferential or tangential view are arranged apparently irregularly or in spirals. But some woods when exam ined from the same view-points show lines running across the grain and producing what is known as ripple-marking: this is caused by either contiguous fibres ending at the same levels with little or no overlapping or medullary rays placed at the same levels, so that the structure of the wood is tiered. Such structure is often shown by true American mahoganies.